The seeds contain an oil rich in lignan, proteins, and the enzyme edestinase (St. Angelo and Ory 1970). The growth hormone zeatin has been found in immature seeds (Rybicka and Engelbrecht 1974). The seeds also contain the alkaloids cannabamine A, B, C, and D, piperidine, trigonelline, and L-(+)-isoleucine-betaine (Bercht et al. 1973). Hemp seed oil, which is obtained by cold pressing the seeds, is very rich in unsaturated fatty acids (vitamin F).
The pollen contains Δ[SUP]9[/SUP]-THC as well as THCA, an alkaloid-like substance, flavones, and phenolic compounds (Paris et al. 1975).
The leaves of Cannabis sativa have be shown to contain choline, trigonelline, muscarine, an unidentified betaine, and, astonishingly, hordenine, a β-phenethylamine alkaloid present in many cacti (El-Feraly and Turner 1975). The leaves of Thai and African populations have yielded water-soluble glycoproteins, serine-O-galactoside, and hydroxyproline (Hillestad and Wold 1977; Hillestad et al. 1977).
The roots of Cannabis sativa have been found to contain friedelin, epifriedelinol, N-(p-hydroxy-β-phenethyl)-p-hydroxy-trans-cinnamamide, choline, and neurine as well as the steroids stigmast-5-en-3β-ol-7-on (= 7-keto-β-sitosterol), campest-5-en-3β-ol-7-on, and stigmast-5,22-dien-3β-ol-7-on (Slatkin et al. 1975).